The game has also another new original character, Chizuru Urashima (voiced by Mariko Kouda).

Making an appearance for the first time are Nintendo characters Chrom and Lucina from Fire Emblem Awakening, and Fiora and Metal Face from Xenoblade Chronicles. The game features pairs of characters from all three participating companies ( Capcom, Namco and Sega), as well as solo characters who can provide assist attacks during combat. Chizuru Urashima and her dimension-hopping vehicle the Dragonturtle Mk.1, travel across worlds and meet allies both old and new, it is revealed that Ouma's plan involves the gathering of the fragments of '99', the final boss from Namco × Capcom, and the mysterious '101 Embryo Plan'. The story begins with Reiji Arisu and Xiaomu traveling to Shibuya to investigate a set of mysterious 'Golden Chains' that have suddenly appeared, and it soon becomes apparent that Ouma is involved when Saya and Sheath (from Ouma's American branch) make their appearance. The story is a continuation of Namco × Capcom's plot, and focus on the conflict between the secret agency Shinra and the chaos-spreading Ouma organization. You can help Capcom Database by expanding it.

In battle, players use well-timed attacks and combos to do as much damage to the opponent as possible before their turn ends. Characters are paired up into 'Pair Units', with 'Solo Units' consisting of a single character able to assist them when linked together.

Like its predecessor, Project × Zone 2 is a tactical role-playing game in which players strategically maneuver their characters across a field to battle with the enemy.

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